Skip to main content

Shane Heschel

Shane Heschel

Associate Professor, Chair

Research Interests

Physiological ecology of plant populations, particularly in stressful environments; local adaptation and the evolution of physiological mechanisms; factors driving the local extinction of plant populations; conservation genetics.

Personal Interests

Hiking, blues guitar, baseball, and pie.


Heschel, M.S., K. Dalton, M. Jamason, A. D'Agnese, and L.G. Ruane. 2017. Drought Response Strategies of Clarika gracilis (Onagraceae) Populations from Serpentine and Nonserpentine Soils. International Journal of Plant Sciences 178.313-319.

Maruyama, C., Z. Goepfert, K. Squires, T. Maclay, Q. Teal-Sullivan, and M.S. Heschel. 2016. Effects of population site and maternal drought on establishment physiology in Impatiens capensis MEERB. (Balsaminaceae). Rhodora 118:32-45.

Craine, E.B., A. Evankow, K.B. Wolfson, K. Dalton, H. Swedlund, C. Bowen, and M.S. Heschel. 2016. Physiological response of Tamarix ramosissima (Tamaricaceae) to a biological control agent. Western North American Naturalist 76:339-351.

Behrman, E.L., S.S. Watson, K.R. O'Brien, M.S. Heschel, and P.S. Schmidt. 2015. Seasonal variation in life history traits in two Drosophila species. Journal of Evolutionary Biology. doi: 10.1111/jeb.12690.

Heschel, M.S., A. Evankow, K.B. Wolfson, J.E. Carlson, and K.E. Holsinger. 2014. Drought response diversification in African Protea species. International Journal of Plant Sciences 175:442-449.

Bibee, K., K. Shishido, R.P. Hathaway, and M.S. Heschel. 2011. Population differentiation of Impatiens capensis (Balsaminaceae) at the range limit. International Journal of Plant Sciences 172:211-219.

Myerburg, M.M., J.J. Ebersole, L.G. Shoemaker, and M.S. Heschel. 2011. Limited plasticity of biomass allocation in alpine forbs, Pikes Peak, Colorado. Western North American Naturalist 71:431–435.

Stinchcombe, J.R., R. Izem, M.S. Heschel, B.V. McGoey, and J. Schmitt. 2010. Across-environment genetic correlations and the frequency of selective environments shape the evolutionary dynamics of growth rate in Impatiens capensis. Evolution 64:2887-2903.

Boggs, J., K. Loewy, K. Bibee, and M.S. Heschel. 2010. Phytochromes influence stomatal conductance plasticity in Arabidopsis thaliana. Plant Growth Regulation 60:77-81.

Lovell, J., J. Gibson, and M.S. Heschel. 2009. Disturbance regime mediates riparian forest dynamics and physiological performance, Arkansas River CO. American Midland Naturalist 162:289-304.

Heschel, M.S., C.M. Butler, D. Barua, G.C.K. Chiang, A. Wheeler, R.A. Sharrock, G.C. Whitelam, and K. Donohue. 2008. New roles of phytochromes during seed germination. International Journal of Plant Sciences 169:531-540.

Riginos, C., M.S. Heschel, and J. Schmitt. 2007. Maternal effects of drought stress and inbreeding in Impatiens capensis (Balsaminaceae). American Journal of Botany 94:1984-1991.

Heschel, M.S., J. Selby, C. Butler, G.C. Whitelam, R.A. Sharrock, and K. Donohue. 2007. A new role for for phytochromes in temperature-dependent germination. New Phytologist - 174:735-741.

Donohue, K., M.S. Heschel, G.C.K. Chiang, C.M. Butler, and D. Barua. 2007. Phytochrome mediates germination responses to multiple seasonal cues. Plant, Cell and Environment 30:202-212

Schmidt, P.S., A.B. Paaby, and M.S. Heschel. 2005. Correlations of reproductive diapause and life history tradeoffs in Drosophila melanogaster. Evolution 59:2616-2625.

Heschel, M.S., N.J. Hausmann, and J. Schmitt. 2005. Testing for stress-dependent inbreeding depression in Impatiens capensis (Balsaminaceae). American Journal of Botany 92:1322-1329.

Heschel, M.S. and C. Riginos. 2005. Mechanisms of selection for drought stress tolerance and avoidance in Impatiens capensis. American Journal of Botany 92:37-44.

Heschel, M.S., S.E. Sultan, S. Glover, and D. Sloan. 2004. Population differentiation and plastic responses to drought stress in the generalist annual, Polygonum persicaria. International Journal of Plant Sciences 165:817-824.

Heschel, M.S., J.R. Stinchcombe, K.E. Holsinger, and J. Schmitt. 2004. Natural selection on light response curve parameters in the herbaceous annual, Impatiens capensis. Oecologia 139:487-494.

Huber, H., N. Kane, M.S. Heschel, J. Banta, A. Leuck, and J. Schmitt. 2004. Frequency and microenvironmental pattern of selection on plastic shade avoidance traits in a natural population of Impatiens capensis. American Naturalist 163:548-563.

Schmitt, J., J.R. Stinchcombe, M.S. Heschel, and H. Huber. 2003. The adaptive evolution of plasticity: phytochrome-mediated shade avoidance responses. Integrative and Comparative Biology 43:459-469.

Heschel, M.S., K. Donohue, N.J. Hausmann, and J. Schmitt. 2002. Population differentiation and natural selection for water-use efficiency in Impatiens capensis (Balsaminaceae). International Journal of Plant Sciences 163:907-912.

Heschel, M.S. and N.J. Hausmann. 2001. Population differentiation for abscisic acid responsiveness in Impatiens capensis (Balsaminaceae). International Journal of Plant Sciences 162:1253-1260.

Donohue, K., E. Hammond Pyle, D. Messiqua, M.S. Heschel, and J. Schmitt. 2001. Adaptive divergence in plasticity in natural populations of Impatiens capensis and its consequences for performance in novel habitats. Evolution 55:692-702.

Donohue, K., D. Messiqua, E. Hammond Pyle, M.S. Heschel, and J. Schmitt. 2000. Evidence of adaptive divergence in plasticity: density- and site- dependent selection on shade avoidance responses in Impatiens capensis. Evolution 54:1956-1968.

Donohue, K., D. Messiqua, E. Hammond Pyle, M.S. Heschel, and J. Schmitt. 2000. Density dependence and population differentiation of genetic architecture in Impatiens capensis in natural environments. Evolution 54:1969-1981.

Paige, K. N. and M. S. Heschel. 1996. Inbreeding depression in scarlet gilia: a reply to Ouborg and Van Groenendael. Conservation Biology 10:1292-1294.

Heschel, M. S. and K. N. Paige. 1995. Inbreeding depression, environmental stress and population size variation in scarlet gilia, Ipomopsis aggregata. Conservation Biology 9:126-133.

Activities & Interests


Drought Response Diversification in African Protea Species

Physiological Response of Tamarix Ramosissima (Tamaricaceae) to a Biological Control Agent

A New Role for Phytochromes in Temperature-Dependent Germination

Mechanisms of Selection for Drought Stress Tolerance & Avoidance in Impatiens Capensis (Balsaminaceae)

Testing for Stress-Dependent Inbreeding Depression in Impatiens Capensis (Balsaminaceae)

Population Differentiation of Impatiens Capensis (Balsaminaceae) at the Range Limit

Disturbance Regime Mediates Riparian Forest Dynamics and Physiological Performance, Arkansas River, CO

Phytochromes Influence Stomatal Conductance Plasticity in Arabidopsis thaliana

Regular Classes

BE105 - Biology of Plants
EV209 - Ecology
BE280 - Population Genetics
BE362 - Desert Plant Ecophysiology
BE365 - Plant Physiology


    1991 BS University of Illinois, Urbana

    1994 MS University of Illinois, Urbana

    2000 PhD Brown University